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Upper Jurassic Pterosaurs

The Upper Jurassic period had a wider range of pterosaurs than any other time.  This period continued until about 136 million years ago when significant changes occurred.  The main United Kingdom Stages are as follows.
Portlandian (south) or
     Volgian (north)
Upper Kimmeridgian
Lower Kimmeridgian
Upper Oxfordian
Lower Oxfordian
Callovian
Lulworth Beds and Lower Purbeck Beds
Portland Beds
Upper Kimmeridge Clay
Lower Kimmeridge Clay with shales
Upper Corallian Beds and Ampthill Clay
Lower Corallian Beds and Upper Oxford Clay
Lower Oxford Clay
Kellaways Beds
Upper Cornbrash
Within the United Kingdom there have been several pterosaur finds,  Rhamphorhynchus from the Oxford Clay,  Pterodactylus and Germanodactylus from the Kimmeridge Clay.  Doratorhynchus, an early Ornithosaur from the Purbeck Beds.  There are also individual bones that are from other species of pterosaur.

The world wide specimens are numerous and show a great deal of information about many species.  This is largely due to a number of fine sites where many pterosaurs have been preserved intact and articulated.  Some sites from this period show limited soft tissue preservation, with wing membranes and body hair intact.

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Anurognathidae

Anurognathus ammoni A rhamphorhynchoid pterosaur with a short snout and a short tail found in the lower Tithonian Lithographic Limestone near Solnhofen, Bavaria, Germany.  Wingspan 50 cm.
Batrachognathus volans Known from two specimens this pterosaur has a distinct short snout and wide jaw.  It comes from the Karatau Upper Jurassic sediments of Kazakhstan.
Dendrorhynchoides curvidentatus Collected from the Zhangijagou locality from the Lower Yixian Formation
Jaholopterus ningchengensis A fine complete specimen showing near complete wing membrane and hair. From the Lower Yixian Formation at Nincheng, Inner Mongolia.  This specimen represents the most complete and largest known Anurognathid.

Azhdarchidae

Doratorhynchus validus A single cervical vertebra from the Purbeck Limestone of Britain.  Jurassic/Cretaceous border sediments.
Un-named Azhdarchid Post cranial skeleton from Solenhofen.

Ctenochasmidae

Beipiaopterus chenianus Partially preserved skeleton from Liaoning Province, China
Ctenochasma elegans A small pterodactyl with long slender teeth.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Ctenochasma gracile Two complete specimens and five fragmentary specimens from the Lower Tithonian, Solnhofen, Lithographic Limestone, Bavaria, Germany.
Ctenochasma porocristata Jaw fragment from the Solnhofen, Lithographic Limestone, showing a porous bony crest on the skull.
Ctenochasma roemeri Lower jaw from the Marine Limestone of Hanover, Lower Saxony, Germany.
Ctenochasma sp. Lower Portlandian, Haut Marne, France.
Ctenchasma taqueti Lower Tithonian of St. Dizier, France
Gnathosaurus subulatus This genus shows many similarities with Ctenochasma though the teeth are less closely packed and there is a distinct saggital crest on the skull.  Lower Tithonian, Solnhofen, Lithographic Limestone, Bavaria, Germany.
Gnathosaurus macrurus A series of vertebrae from the Purbeck Limestone of the United Kingdom.
Huanhepterus quingyangensis This pterosaur is similar to Gmathosaurus, though having a long neck.  Ordos Basin, China.
Plataleorhynchus streptophorodon Fossil of a pterosaur snout with a terminal spatulate shape.  Purbeck Limestone, Dorset, England.

Dimorphodontidae

Dimorphodon weintraubi Finely preserved partial skeleton showing primitive foot structure.  callovian, Tamaulipas, Mexico.  Similarities with Dimorphodon macronyx.

Dsungeripteridae

Dsungaripterus brancai Obere Saurier-Mergel, Tendaguru, Tanzania.
Dsungeripterus sp. Partial skeleton from the Kimmeridgian of Germany

Gallodactylidae

Gallodactylus canjuersensis Specimen from Canjuers in Southern France.
Cycnorhamphus suevicus Well documented specimen from the upper Jurassic Limestone of Wuttemberg, Bavaria.
Feilongus youngi Jianshangou Bed, lower Yixian Formation at Heitizigou in Beipiao, Liaoning Province, China

Germanodactylidae

Germanodactylus cristatus Specimen from the Lower Tithonian, Solnhofen, Lithographic Limestone, Bavaria, Germany.
Germanodactylus rhamphastinus Specimen from the Momsheim Limestone, Bavaria, slightly more recent than the Solnhofen specimen.
Germanodactylus sp. Wing and leg fragments from the Kimmeridge Clay, Dorset, England.
Tendaguripterus recki A pterosaur jaw from Tanzania

Pterodactylidae

Diopecephalus longicollum A large pterodactyl with a wingspan of 1.45 metres.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Eosipterus yangi Articulated wings and legs of a medium sized pterodactyle.
Pterodactylus antiquus Several specimens of this pterodactyl are known from the Lower Tithonian, Lithographic Limestone, Bavaria, Germany.  This was the first pterosaur to be described by Cuvier in 1784.
Pterodactylus grandis One of the largest Jurassic pterodactyls with a wingspan of 2.4 metres.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Pterodactylus grandipelvis Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Pterodactylus kochi The most common of the fossil pterodactyls, many fine specimens have been found, including juvenile remains.  Adults have a wingspan of 50 cm.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Pterodactylus micronyx Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Pterodactylus cerinensis This pterodactyl was described from a single humerus.  A tibia was also assigned to this species.  Kimmeridgian, Lithographic Limestone, Cern, Department Ain, France.
Pterodactylus suprajurensis Individual remains of a large pterosaur were assigned together to this species.  Purbeckian, Boulogne-sur-Mer, France.
Pterodactylus manseli Individual bones of a small pterodactyl are grouped together.  Kimmeridge Clay, Weymouth, Dorset, England.
Pterodactylus pleydelli a small number of similar individual bones from the Kimmeridge Clay, Weymouth, Dorset, England.
Pterodactylus arningi Fragmentary remains, Obere Saurier-Mergel, Tendaguru, Tanzania.
Pterodactylus maximus Fragmentary remains, Obere Saurier-Mergel, Tendaguru, Tanzania.

Rhamphorhynchidae

Rhamphorhynchus gemmingi Over eight fine specimens and other fragmentary remains.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Rhamphorhynchus intermedius At least ten specimens assigned to this species.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Rhamphorhynchus longicaudus In excess of twelve specimens of this pterosaur from the Lower Tithonian, Lithographic Limestone, Bavaria, Germany.  many more specimens showing incomplete remains.
Rhamphorhynchus longiceps Six fine specimens including one near complete animal.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Rhamphorhynchus muensteri About forty specimens assigned to this species.  Some very fine examples including wing membrane preservation.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Rhamphorhynchus jessoni Fragmentary remains of wing bones and joints.  Oxfordian, Oxford Clay, England.
Rhamphorhynchus tendagurensis Upper saurier-marl, Tendaguru, Tanzania.
Rhamphorhynchus sp. Upper Callovian, Pedroga, Lower Kimmeridgian, Guimarota near Leiria, Portugal.
Cacibupteryx carabensis Skull and partial wing from the Jagua Formation, Cuba.
Odontorhynchus aculeatus Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Pterorhynchus wellnhoferi Rhamphorhynchoid exhibiting a head crest.
Scaphognathus crassirostris There are at least five specimens of this sturdy pterosaur and many fragmentary remains.  Lower Tithonian, Lithographic Limestone, Bavaria, Germany.
Harpactognathus gentryii Jaw fragment from the Morrison Formation, Utah.
Sordes pilosus Two specimens showing wing membrane and body hair preservation.  Upper Jurassic, Karatau, Kazakhstan.

Unclassified

Kepodactylus insperatus Short spined vertebra, left humerus and fragmentary wing bones from Colorado, U.S.A. of Kimmeridgian Age
Dermodactylus montanus Fragment of wing metacarpal from Como Bluff, Wyoming, U.S.A.  This was the first fossil pterosaur to be found in the Americas.
Wyomingopteryx ? Undescribed pterosaur from the U.S.A.  (awaiting details)
Comodactylus ostromi A wing-metacarpal from the Morrison Formation, Como Bluff, Wyoming, U.S.A.  This specimen is of a Rhamphorhynchoid species
Laopteryx priscum Partial skull previously classified as a bird, re-evaluated as a pterosaur.
Mesadactylus ornithosphyos A group of bones associated in a partial, post cranial skeleton, Como Bluff, Wyoming, U.S.A.
Nesodactylus hesperius Post cranial and disarticulated skeleton of a small pterosaur from the Oxfordian of Vinales, Pinar del Rio, Cuba. This specimen is of a Rhamphorhynchoid species
Herbstosaurus pigmaeus pelvis, femur and other fragments of a small pterosaur from the Callovian, Lotena Formation, Neuquen, North Patagonia, Argentina.  probably of a Rhamphorhynchoid species.
Normannognathus wellnhoferi Jaw fragments from Cap de le Heve, Normandy, France.  Upper jaw 6cm and lower jaw 5cm showing several tooth sockets.  Possibly an Ornithocheirid.
Pterosaur sp. Specimen from Solothurn, Switzerland
Utahdactylus keteae Pterosaur remains from the Morrison formation, USA.
Rhamphorhynchinae indet. Wing phalange shaft from Switzerland

Ichnites

Petrachnus saltwashiensis Salt Wash Member, Morrison Formation, Carrizo Mountains, Arizona, USA.
Stokes, W. L. 1957. Pterodactyl tracks from the Morrison Formation. Journal of Paleontology, 31:952–954.
Pteraichnus stokesi Alcova Reservoir region, Casper, Wyoming, USA.
Logue, T. J. 1994. Alcova, Wyoming tracks of Pteraichnus saltwashensis, made
by pterosaurs. Geological Society of America Abstracts with Programs, 26:10.
Connely M. V., 2004, Alcova's Pteraichnus Lagerstatten, 2004 Denver Annual Meeting (7-10th November 2004) Geological Society of America Abstracts with Programs vol.36 No.5 p.67
Pteraichnus longipodus Villar de Rio, Oncala Group, Upper Jurassic boundary, Spain.
Pteraichnus sp. Wierzbica Quarry, Radom, Holy Cross Mountains, Poland.
Pieńkowski G. and NiedŸwiedzki G. 2005; Pterosaur tracks from the early Kimmeridgian intertidal deposits of Wierzbica, Poland. Geol. Quart., 49 (3): 339–346. Warszawa.
Pteraichnus sp. Summerville Formation, Ferron, Utah, USA.
Pteraichnus palacieisaenzi Sundance Formation, Bighorn Canyon National Recreational Area, Wyoming, USA.
Arribas, C. P. and Pérez, E. S. 2000. Huellas de pterosaurios en el Grupo On-
cala (Soria, España). Pteraichnus palaciei-saenzi, nov. ichnosp. Estudios
Geologicos, 56:73–100.