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Upper Cretaceous Pterosaurs

The Upper Cretaceous period was the last of the Mesozoic ages, terminating in a mass extinction at about 65 million years ago.  This age saw the largest of the pterosaurs and some species have been found world wide.  The United Kingdom main stages for this period are as follows.

Maastrichtian

Senonian

Turonian

Upper Cenomanian

Middle Cenomanian

Lower Cenomanian

Upper Chalk and Trimmingham Chalk (Lower Maastrichtian)

Upper Chalk and Norwich Chalk

Middle Chalk

Plenus Marls and basal Melbourne Rock

Lower Chalk and Chalk Basement Beds

Chalk Marl and various local beds

The Upper Cretaceous Stages in the United Kingdom are essentially the Chalk of Southern England.  There are a good variety of pterosaur remains of species like Ornithocheirus and Ornithostoma.  Other species are known from fragmentary evidence.

World wide, the large pterosaurs like the Pteranodontids and Azhdarchids  are widely found along with many other smaller species.

Azhdarchidae

Arambourginiana philadelphidae	Single large cervical vertebra (60 cm) from Jordan
Aralazhdarcho bostobensis
Azhdarcho longicollis	Specimens from Uzbekistan, Central Asia
Azhdarcho sp.	Associated bones from Alberta Canada.  Predator damaged.
Azhdarchid from Solana	Fragmentary vertebra from La Solana, Spain
Azhdarchid from Valencia	Partial vertebrae from Valencia, Spain
Azhdarchid from Hell Creek	Single cervical vertebra in association with a tyrannosaurid.
Bakonydraco galaczi	Mandible and vertebrae from the Bakony Montains, Hungary
Bogolubovia orientalis	fragmentary remains from Saratov, Petrovsk, 'USSR'
Hatzegopteryx thambema	Large pterosaur of Maastrichtian age from the Densus-Ciula Formation, Romania.  Estimated 12 metre wingspan
Montanazhdarcho minor	fragmentary remains of humerus, radius and complete distal carpal of a very large pterosaur
Phosphatodraco mauritanicus	5 cervical vertebrae from the Oulad Abdoun Basin near Khouribga
Quetzalcoatlus northropi	Very large pterosaurs from the Javelin Formation, Big Bend National Park, West Texas, U.S.A.

Nyctosauridae

Nyctosaurus gracilis	Complete Specimen, Niobara Chalk, Santonian, West Kansas, U.S.A.
Nyctosaurus nanus	O. C. Marsh 1881
Nyctosaurus lamegoi	Gramame Formation, Paraiba, Brazil
Nyctosaurus bonneri	A substantial specimen of a large juvenile
Nyctosaurus sp.	Unusual species with a very large head crest and sail.
Muzquizopteryx coahuilensis	Near complete specimen from Mexico

Ornithosauria

Lonchodectes compressirostris	Jaw from the Turonian, Middle Chalk of Burnham, Kent, England
Lonchodectes giganteus	Jaw fragment from the Cenomanian, Lower Chalk, Burnham, Kent, England.  Similar jaw fragments have been found in the Cambridge Greensand
Ornithocheirus cuvieri	distal jaw of a large pterosaur found in the Burnham Chalk Pit, Kent, England
Ornithocheirus fittoni	Jaw fragments from the basal sediments of the Upper Cretaceous.  This species is predominantly a Cambridge Greensand pterosaur from the Lower Cretaceous
Ornithocheirus hlavatschi	Associated wing bones from Czechoslovakia.

Pteranodontidae

Ornithostoma sedgwicki	Fragmentary remains from the Cenomanian, England
Pteranodon species	An extremely popular pterosaur in the perception of the public.
Pteranodon longiceps	Several fine specimens including a crested skull of 1.79 metres in length.  This pterosaur is well documented from the Niobara Chalk of Kansas, U.S.A.
Pteranodon sternbergi	Large Pteranodon with a 1.2 metre jaw and a large vertical skull crest.  Estimated wing span of 7 metres.  Niobara Chalk of Kansas, U.S.A.
Pteranodon sp	Hokkaido, Japan
Pteranodon sp	Pterosaur from The Lebanon

Track ways and Ichnites

Haenamichnus uhangriensis	Large trackway from Korea. Hwang K.-G., Huh M., Lockley M. G., Unwin D. M. and Wright J. L., 2002, New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea, Geological Magazine, 139: 421-435, Cambridge University Press
Pteraichnus sp.	Blackhawk Formation, Utah, USA. Lockley, M. G. 1999. Pterosaur and bird tracks from a new Late Cretaceous locality in Utah. In Gillette, D. D. (ed.). Vertebrate Paleontology in Utah. Utah Geological Survey, Utah Geological Survey Miscellaneous Publication, 99-1:355–359.
Agadirichnus elegans	Maastrichtian, Morocco. Ambroggi, R. et Lapparent, A. F. de, 1954. Découverte d'empreintes de pas de Reptiles dans le Maestrichtien d'Agadir.Notes du Service géologique du Maroc, 10,43-57.