Cearadactylus atrox

This pterosaur is known from a single skull of about 57cm in length from the Santana Formation, north eastern Brazil. The jaw is articulated and uncrushed, showing the position in life. The systematic relationship of this species is unclear and several attempts to clarify this have produced conflicting analyses.
Holotype: F-PV-93 ; Extensive partial skull and mandible
Referred Specimens: GIUA M. 4895; Pair of associated cervical vertebrae.
Unwin, D. M. 2002, On the systematic relationships of Cearadactylus atrox, an enigmatic Early Cretaceous pterosaur from the Santana Formation of Brazil. Mitteilungen Museum für Naturkunde Berlin, Geowissenschaftlichen Reihe, 5, 239-263.